Pages
▼
▼
Tuesday, December 30, 2014
Monday, December 29, 2014
Common Bullfrog (Kaloula taprobanica)
A common frog found in anthropological habitats, paddy fields, plantations and forest borders in both wet and dry lowlands up to about 500m a.s.l. Nocturnal in habits and stay hide under decaying logs, rock crevices, under leaf litter and inside humus or soil during the day time. Its calls resembles the bellowing of a bull.
Saturday, December 27, 2014
Friday, December 26, 2014
Double Branded Crow (Euploea sylvester montana)
Wednesday, December 24, 2014
Tuesday, December 23, 2014
Sunday, December 21, 2014
Saturday, December 20, 2014
Vagrant Finches and Buntings (Family: Fringillidae and Emberizidae) recorded in Sri Lanka
Birds that appear outside their normal range are known as vagrants. This post summarizes up to date published sight records of 3 vagrants of the family Emberizidae (Buntings) and single vagrant of the family Fringillidae (Finches) in Sri Lanka. All four are here considered as unconfirmed vagrants*.
*Species for which there are only one or two sight records exist categorized here as unconfirmed vagrants. Problematic records without sufficient details are also included.
1) Common Rosefinch (Carpodacus erythrinus)
Single sight record at Horton Plains National Park in February 2002 by Uditha Hettige (Siriwardana, U., 2003).
2) Grey-necked Bunting (Emberiza buchanani)
C.D. Kaluthota, Dammithra Samarasinghe, Dilshan de Silva and Hasith de Silva observed and photographed a bird at Nirmalapura, Daluwa area of Kalpitiya peninsula on 16th December 2008. It is the first and only known occurrence of Grey-necked Bunting in Sri Lanka (Kaluthota et. al., 2009)
3) Black-headed Bunting (Emberiza melanocephala)
Single sight record from Udawalawe National
Park in February 2005 by Deepal
Warakagoda is the first record of its occurrence in Sri Lanka (Sirivardana
U. & Warakagoda D. , 2006). Second record of it is also given by Warakagoda D. & Sirivardana U. in/near Yala Block 1 in the report of Ceylon Bird Club for 2008-2011 periods (Warakagoda D. & Sirivardana U., 2011)
4) Red-headed Bunting (Emberiza bruniceps)
Single sight record from Udawalawe National
Park in February 2005 by Deepal
Warakagoda is the only record of its occurrence in Sri Lanka (Sirivardana
U. & Warakagoda D. , 2006)
References:
Kaluthota, C.D., Samarasinghe, D., de Silva, D. & de Silva, H. 2009 First record of the Grey-necked Bunting Emberiza buchanani from Sri Lanka. Malkoha 30(1):4
Siriwardana, U., 2003. Report from the Ceylon Bird Club for 2002. Loris, 23(3&4), 36-39.
Sirivardana U. & Warakagoda D. ,
2006. Report from the
Ceylon Bird Club for 2005 Loris 24(3& 4), 24 - 28
Warakagoda D. &
Sirivardana U., 2011. Report from the Ceylon Bird Club for 2008-2011, Loris 26
(1&2)
Thursday, December 18, 2014
Wednesday, December 17, 2014
Small Grass Yellow (Eurema brigitta rubella)
Locally common butterfly occurs in grasslands of the hills from 200-2200m asl. Its larva feeds on Chamaecrista leschenaultiana, Chamaecrista kleinii , Chamaecrista mimosoides and Chamaecrista nictitans (All Chamaecrista species are known as බිං සියඹලා in Sinhala)
Monday, December 15, 2014
Sunday, December 14, 2014
Bar-winged Flycatcher-Shrike (Hemipus picatus)
An uncommon breeding resident inhabiting forest and well wooded areas of all climatic zones. But much common in the hills and more or less restricted to the riverine forests in the dry zone. It keeps in pairs or groups and often goes with 'Bird waves' of many species. Bar-winged Flycatcher-Shrike feeds on insects in the canopy. It frequently darting out like 'Flycatchers' from its perch to catch flying insects. The breeding season lasts from March to May. The nest is a shallow cup made out of fibres, cobwebs, etc., well camouflages outside with fragments of bark, lichens etc. and places on the upper side of a branch high in the canopy.
සිංහලෙන් කියවන්න >>
Friday, December 12, 2014
Thursday, December 11, 2014
Common Mime (Papilio clytia lankeswara)
Tuesday, December 9, 2014
Monday, December 8, 2014
Vagrant Rails and Crakes (Family: Rallidae) recorded in Sri Lanka
Birds that appear outside their normal range are known as vagrants. This post summarizes up to date published sight records of vagrants of the family Rallidae (Rails and Crakes) in Sri Lanka.
1) Water rail (Rallus aquaticus) R.Indicus
3 or 4 specimens procured by Lieut. Long of H.M. Ceylon rifle regiment at Jayelle (Sic = Jaela) paddy-fields and given to Layard is the only record of the occurrence of this rail in Sri Lanka (Legge V. 1880: 1018).
2) Corn Crake (Crex crex)
Two immature females procured from Bambalapitiya, one by Dr. Thelma Gunawardane from a garden near the sea on 9th October 1970 and another one in exhausted condition by David Bartholomeusz in the same area on 18th September 1972 are the only records of Corn Crake in Sri Lanka. Both are believe to be just arrived birds from the Indian mainland (Phillips W.W.A., 1978:23). Another previous sight record exists in 1950 on board a ship close to Ceylon. (Hoffmann, T. W., 1974)
References:
Hoffmann, T. W., 1974. Notes From The Ceylon Bird Club 1970-1973. Loris, 13(4), 211-214.
Legge V., 1880. A History of the birds of Ceylon 1983 second edition.
Phillips W.W.A., 1978 Annotated checklist of the Birds of Ceylon (Sri Lanka) 1978 revised edition.
Friday, December 5, 2014
Thursday, December 4, 2014
Wednesday, December 3, 2014
Monday, December 1, 2014
Dingy Lineblue (Petrelaea dana)
* Terminalia arjuna (කුඹුක්) is also known as one of its larval host plant.
Sunday, November 30, 2014
Friday, November 28, 2014
Vagrant Swifts (Family: Apodidae) recorded in Sri Lanka
Birds that appear outside their normal range are known as vagrants. This post summarizes up to date published sight records of four unconfirmed vagrants* of the family Apodidae (Swifts ) in Sri Lanka.
*Species for which there are only one or two sight records exist categorized here as unconfirmed vagrants. Problematic records without sufficient details are also included.
1) White-throated Needletail (Hirundapus caudacutus)
1) White-throated Needletail (Hirundapus caudacutus)
Amila Salgado reported a sight record of White-throated Needletail first time in Sri Lanka in Pokunutenna (located in the North-Eastern boundary of the Udawalawe national park) on 22nd February 2007. He had observed several of them in a flock of swifts which included little swifts, Indian swiftlets and Asian palm swifts. He has also mentioned about a sight record of this swift in 2003 at Sinharaja by a visiting bird watcher though which was not reported formerly (Salgado A., 2007). A sight record of two White-throated Needletails on 19th February 2003 in Sinharaja is mentioned under ‘List of rejections (after 1990)’ of Ceylon bird club (http://www.ceylonbirdclub.org/List-of-Rejections.pdf). This is probably the same sight record Salgado was referring above.
2) Pallid Swift (Apus pallidus)
A swift probably of this species was photographed in South-east Sri Lanka (Rasmussen P.C. and Anderton J.C., 2012: 259-260)
3) Fork-tailed Swift [Pacific Swift/Large White-rumped swift] (Apus pacificus)
First record in Sri Lanka by Deepal Warakagoda from Bundala national Park in 28th November 1996 (Hoffmann, T. W., 1997 & Warakagoda, D., 1999). A sight record by M.J. Pointon (UK) at Bentota in December 1997 is the second record of it in Sri Lanka (Hoffmann, T. W., 1999). However another record is given as the second
report of it in/near Yala Block 1 by Warakagoda D. & Sirivardana
U. in the summery of bird club notes for 2008-2010 periods (Warakagoda D.
& Sirivardana U., 2011)
4) Dark-rumped Swift (Apus acuticauda)
A sight record at Kandy in February 1993 by A.J. Vincent is the only record (Kotagama S. & Ratnavira G., 2010: 257)
References:
Hoffmann, T. W., 1999. Ceylon Bird Club Notes, 1997. Loris, 22(1), 57-58.
Hoffmann, T. W., 1997. Ceylon Bird Club Notes 1996. Loris, 21(3), 112-113.
Kotagama, S., Ratnavira, G. 2010. An illustrated Guide to the Birds of Sri Lanka. Field Ornithology Group of Sri Lanka, Colombo.
Rasmussen P.C. and Anderton J.C., 2012. Birds of South Asia - The Repley Guide Volume 2 Attributes and Status, second edition.
Salgado A., 2007. A sight record of White-throated Needletail Hirundapus caudacutus in Pokunutenna, Sri Lanka Siyoth 2(2): 43.
Warakagoda, D., 1999. Ceylon Bird Club Notes,1998. Loris, 22(2), 33-34.
References:
Hoffmann, T. W., 1999. Ceylon Bird Club Notes, 1997. Loris, 22(1), 57-58.
Hoffmann, T. W., 1997. Ceylon Bird Club Notes 1996. Loris, 21(3), 112-113.
Kotagama, S., Ratnavira, G. 2010. An illustrated Guide to the Birds of Sri Lanka. Field Ornithology Group of Sri Lanka, Colombo.
Rasmussen P.C. and Anderton J.C., 2012. Birds of South Asia - The Repley Guide Volume 2 Attributes and Status, second edition.
Salgado A., 2007. A sight record of White-throated Needletail Hirundapus caudacutus in Pokunutenna, Sri Lanka Siyoth 2(2): 43.
Warakagoda, D., 1999. Ceylon Bird Club Notes,1998. Loris, 22(2), 33-34.
Warakagoda D. &
Sirivardana U., 2011. Report from the Ceylon Bird Club for 2008-2011, Loris 26
(1&2)
Monday, November 24, 2014
Tamil Bushbrown (Mycalesis subdita)
Tamil Bushbrown is an endemic butterfly occurs in scrublands, home gardens and grasslands in dry and intermediate zones below 750m elevation. It is fond of rotting fruits. Larva feeds on Ischaemum timorense in the field and fed on Panicum maximum(Guinea Grass/රට තණ) and Axonopus compressus (Carpet Grass/පොටු තණ) in the lab when offered.
Sunday, November 23, 2014
Friday, November 21, 2014
Possible range extension of Rasboroides nigromaginatus(?)
Note : - Sudasinghe et al., 2018 based on a phylogenetic study, resolved the issues raised in this post at the end. Now Rasboroides nigromarginatus is considered as a synonym of Rasboroides vaterifloris
Sudasinghe, H., Herath, J., Pethiyagoda, R. and Meegaskumbura, M. , 2018, ‘Undocumented translocations spawn taxonomic inflation in Sri Lankan fire rasboras (Actinopterygii, Cyprinidae)’. PeerJ [Online journal], (20 December 2018). Link >>
Endemic genus Rasboroides is restricted to the shaded rain
forest streams of South-western wet zone from Kalu to Walawe drainages. A single species of Rasboroides formerly
known as Rasbora vaterifloris was divided into four different species
recently base on morphological differences. Table 1 summarized some of their
morphological features with distribution in the country base on Batuwita S. et
al., 2013.
Rasboroides
nigromarginatus ( Black Line Golden Rasbora/කලු-ඉරි හල්මල්දන්ඩියා[Kaluiri Halmaldandiya])
Rasboroides nigromarginatus was
first described by a German scientist H. Meinken in 1957 as a new species base on a single specimen
collected from the ‘Ceylon’ without knowing exact locality. He had found that specimen among some exported ornamental fishes from Sri Lanka and considered it as a
synonymy of Rasboroides vaterfloris (Deraniyagala,
1958 & Pethiyagoda, 1991). It has not been collected since then until the rediscovery in 1994 from Athwelthota by R. Pethiyagoda & K. Manamendra Arachchi and
independently in 2010 by members of Wildlife
Conservation Society, Galle.
With the recent review of Rasboroides group it is now considered as a valid species occurs only in Athwelthota area of Kalu
river basin. Here we record a possible range extension of this species again on
Kalu river basin but considerable
distance away from the type locality. We
would like to keep exact locality undisclosed, considering past
experiences of extinction of some rare
endemic fish species from certain localities without proper protection, due to
excess collection for export and local ornamental fish industry (Ex. Extinction
of bright color Rasboroides species from Puwakpitiya Dola in Indikada Mookalana and total extinction of them from Madakada Mookalana - Chamikara S., 2012).
We were able to observe and photograph four live specimens of
Rasboroides in a tributary of Kalu
River in the Kiriella area of the Rathnapura district. The stream is bordered to a rubber plantation
but with well wooded banks with several indigenous plants. One specimen we observed was bit bigger than other three and lack black color of un-branched
ray of dorsal and other fins on the leading edge (However it is slightly
visible on the dorsal fin ray on nearly half of its length – See picture). But three small specimens had prominent black
lines along leading edge of all their fins. Other than that large specimen was more yellowish on its dorsal and upper half of the side and fins while small
specimens had more silver in dorsal as well as venter and sides. However orange red color of dorsal fin and
orange of caudal fin was not observed in any specimens. Taking accurate
measurements of the specimens was impossible with living fishes due to obvious
reason. But approximate size of the large specimen was visually estimated as
30mm and about 20mm of small three. Body
depth of all 4 specimens was around 30% SL.
Three out of four species of
Rasboroides have been recorded from Kalu
river basin and R. vaterifloris reported from Small shaded streams and rivulets
in and around Gilimale is the closet
Rasboroides locality recorded so far to the locality of the current specimens. However
morphological features of them much resembles to the R. nigromaginatus hitherto
only recorded from Athwelthota
area.
Is this a hybridization of R. nigromaginatus and R. vaterifloris? Why only small individuals have prominent black ray on leading edges of fins? Are they juveniles and would it be faded away in adult specimens? Or we are here dealing with two different species (We have observed that small individuals are schooling separately during our observation)? More research need to be done before come to a conclusion especially with exact measurements and scale counts with preserved specimens. But which is beyond our scope and for the time being we would like just to report this observation and highlight the need of conducting more research on our ichthyofauna and also most importantly conserving of their habitats with special consideration towards the streams and rivulets outside the protected forest reserves in Southwestern wet zone of the country. Because those water sources are highly vulnerable to the human activities (We got to known that excessive fishing have been practicing in above area some time even with using explosives which destroy every living inhabitants of the stream from big to small). There are enough possibilities of discovering new species not only of fishes but also other animals as well in these unprotected areas and it is needed to conserve them before those ‘to be discovered’ species would be vanishing from the world before even known to the scientific community.
Is this a hybridization of R. nigromaginatus and R. vaterifloris? Why only small individuals have prominent black ray on leading edges of fins? Are they juveniles and would it be faded away in adult specimens? Or we are here dealing with two different species (We have observed that small individuals are schooling separately during our observation)? More research need to be done before come to a conclusion especially with exact measurements and scale counts with preserved specimens. But which is beyond our scope and for the time being we would like just to report this observation and highlight the need of conducting more research on our ichthyofauna and also most importantly conserving of their habitats with special consideration towards the streams and rivulets outside the protected forest reserves in Southwestern wet zone of the country. Because those water sources are highly vulnerable to the human activities (We got to known that excessive fishing have been practicing in above area some time even with using explosives which destroy every living inhabitants of the stream from big to small). There are enough possibilities of discovering new species not only of fishes but also other animals as well in these unprotected areas and it is needed to conserve them before those ‘to be discovered’ species would be vanishing from the world before even known to the scientific community.
The Habitat of Rasboroides nigromaginatus(?)
R.
vaterifloris
|
R.
nigromaginatus
|
R. pallidus
|
R. rohani
|
|
Male –Upper
body color
|
Golden brown
lightening on side to silvery, scattered with melanophores
|
Golden
orange, sides and venter silver
|
Dorsal color
varies from pinkish-grey to dark golden yellow, lighter latterly. Silver on
venter
|
Dorsally
greyish-yellow, golden yellow on sides and silvery-white on venter.
|
Male- belly
|
Silver
|
Silver
|
Silver
|
|
Dorsal Fin
|
Hyaline with
scattered melanophores
|
Orange red, Longest
un-branched dorsal fin ray black along its entire length.
|
Golden yellow
|
Golden yellow
|
Anal fin
|
Hyaline with
scattered melanophores
|
Yellow with
scattered melanophores. Unbranched rays of leading edge black
|
Golden yellow
|
Yellow-orange
|
Pectoral fin
|
Hyaline with
scattered melanophores
|
Yellow with
scattered melanophores. Unbranched rays of leading edge black
|
Golden yellow
|
Golden yellow
|
Pelvic fin
|
Hyaline with
scattered melanophores
|
Yellow with
scattered melanophores. Unbranched rays of leading edge black
|
Golden yellow
|
Golden yellow
|
Caudal fin
|
Greenish
Yellow
|
Orange.
|
Lighter with
upper lobe hyaline
|
Yellow-orange
on lower lobe. Upper lobe hyaline.
|
Sclera
|
Upper third
of sclera deep Orange
|
Dense melanophores
on sclera above and below giving effect of black bar on eye (Bothe M and F)
|
||
Opercle
|
Suffused with
dark red (Bothe M and F)
|
|||
Female
|
Similar to
male but lighter in color, yellowish tan dorsally whitish with scattered
melanophores on sides, belly white
|
Similar but
lighter in color. Lacking black dorsal and anal fin margins and yellowish tan
dorsally. Whitish with scattered melanophores on sides, , belly white
|
Overall light
whitish orange. Lighter ventrally and on fins, which are partly hyaline
|
Sexes alike
|
Fins
|
Pectoral fin
reaching beyond pelvic fin base; pelvic fin reaching beyond anal fin origin.
Dorsal fin origin located halfway between snout-tip and hypural notch.
|
Pectoral fin
reaching beyond base of pelvic fin; pelvic fin reaching beyond origin of anal
fin; Dorsal fin origin located halfway between snout-tip and hypural notch.
|
Pectoral fin
reaching base of pelvic fin; pelvic fin reaching origin of anal fin. Dorasl
fin origin located halfway between snout-tip and hypural notch.
|
Pectoral fin
reaching base of pelvic fin; pelvic fin reaching origin of anal fin. Dorasl
fin origin located halfway between snout-tip and hypural notch.
|
Dorsal fin
origin
|
1 scale width
behind the pelvic fin origin
|
1 scale width
behind the pelvic fin origin
|
3 scale width
behind the pelvic fin origin
|
3 scale width
behind the pelvic fin origin
|
Pelvic fin
origin
|
Midway
between pectoral fin base and anal fin origin
|
Closer to
anal fin origin than to pectoral fin base.
|
Falling
midway between pectoral fin base and anal fin origin
|
Closer to
anal fin origin than to pectoral fin base
|
Dorsal margin
of the operculum
|
More or less
straight
|
Deep
indentation
|
||
Dorsal
profile of the head
|
Approximately
straight behind the level of the eye in the male
|
Slightly
concave
|
Slightly
concave behind level of eye, arched thereafter
|
Approximately
straight posterior to level of eye
|
Male Interorbital
width
|
29-33 % HL
|
17-22 % HL
|
||
Male Internarial
width
|
19-21 % HL
|
11-17 % HL
|
||
Female anal
fin depth
|
23.5-24.8 % SL
|
17.4 – 22.7 %
SL
|
||
Female interorbital
width
|
25-33 % HL
|
16-21 % HL
|
33-36%HL
|
25-28 % HL
|
Female Caudal-peduncle depth
|
12.4 – 13.4 %
SL
|
10.4 – 12.3 %
SL
|
||
Female Caudal-peduncle length
|
18.6-21.2% SL
|
21.1-25 % SL
|
||
Body depth
|
31-32.8 % SL
( 31.1-32.8 in males and 31 – 31.7 in females)
|
26.9-33% SL
(28.2-33 in males and 26.9-29.1 in females)
|
34.3-37.5%SL
of males and 33-36.7in females
|
35.6 -40% SL
in males and 33.3-36.8 % SL in females
|
Size
|
Male –
23.9-29.8mm Female – 22.1 – 25.5mm
|
Males 26.2 –
30.2 Female – 26.5 – 27.3mm
|
Up to 24.6 mm
SL (males 21.5-24.6 and females 20.2- 20.7)
|
Up to 35.5mm
SL (Males 25.3-35.5 and Females 23-30.8)
|
Scales on
transverse line on the body
|
½ 8 ½
|
½ 8 ½
|
½ 6 ½ or ½ 7
½ ( ½ 6[3] or 7 ½ [12] )
|
½ 8 ½
|
Scales on caudal
peduncle
|
½ 4 ½
|
½ 4 ½
|
½ 3 ½
|
½ 4 ½
|
Predorsal
scales
|
10[2],11[6],12[2]
|
11[4] or
12[6]
|
10[7],11[5],12[3]
|
11[5],12[6],13[4]
|
Prevental
scales
|
15[2] or
16[8]
|
14[2], 15[7],
16[3]
|
13[7],14[4],15[2]
|
14[3],15[9],16[3]
|
Abdominal
vertebrae
|
12[1] or
13[1]
|
12
|
11[4], 12[1],
13[1]
|
12
|
Caudal
vertebrae
|
16[1] or
17[1]
|
16
|
16[2] or
17[4]
|
17
|
Total
vertebrae
|
29
|
28
|
28-29
|
29
|
Scales on
lateral series on body
|
21[2],22[1],23[1],24[5],25[1]
|
23[1],24[1],25[7],26[2],27[1]
|
20[1], 21[1], 22[7],23[0],24[6]
|
25[3],26[8],27[1],28[3]
|
Perforated
scales arterially
|
4[3],5[2],6[5]
|
1[1],2[5],3[7]
|
1[11] or 2[4]
|
1[6],2[3],3[5]
|
Lateral
processes to the kinethmoid
|
2
|
1
|
||
Female eye
diameter
|
39 – 40% HL
|
29-34 % HL
|
||
Pharyngeal-teeth
formula
|
5,4,3-3,4,5
|
5,4,3-3,4,5
|
5,4,3-3,4,5
|
5,4,2 -2,4,5
|
Dorsal fin
rays
|
3 unbranched
and 7 ½ bracnched
|
3 unbranched
and 7 ½ bracnched
|
3 unbranched
and 7 ½ bracnched
|
3 unbranched
and 7 ½ bracnched
|
Anal fin rays
|
3 unbranched
and 5 ½ [2] or 6 ½[8] branched
|
3 unbranched and
6 ½ branched
|
3 unbranched
and 6 ½ branched
|
3 unbranched
and 6 ½ branched
|
Caudal fin
rays
|
17 branched
rays 9 on upper and 8 on lower lobes
|
17 branched
rays 9 on upper and 8 on lower lobes
|
17 branched
rays 9 on upper and 8 on lower lobes
|
17 branched
rays 9 on upper and 8 on lower lobes
|
Pectoral fin
rays
|
1 unbranched
and 11 ½ [4] or 12 ½ [6] branched
|
1 unbranched
and 11 ½ branched
|
1 unbranched
and 11 ½ branched
|
1 unbranched
and 11 ½ branched
|
Pelvic fin
rays
|
1 unbranched
and 6 ½ [3] or 7 ½ [7] branched
|
1 unbranched
and 6 ½ branched
|
1 unbranched
and 6 ½ branched
|
1 unbranched
and 6 ½ branched
|
Lateral line
|
Incomplete
|
Incomplete
|
Incomplete
|
Incomplete
|
Distribution
|
Restricted to
Kalu basin. Small shaded streams and rivulets in and around Gilimale. Type
locality – Ilukwaththa ferry of Kalu river at Gilimale
|
Restricted to
the Athwelthota of Kalu basin
|
Shallow slow
flowing densely shaded low land rain forest streams with sandy silt
substrates of Kalu (Mahakalupahana, Yagirala), Benthara(Bambaravana), Gin
(Kottawa-Kombala FR)Polathu-Modara (Kottawa-Kombala FR), Nilvala (Dediyagala
FR) basins.
|
Suriyakanda
in Walawe river basin. Shallow slow flowing shady streams with a sandy-silt
substrates and dense leaf litter in submontane forests.
|
Literature Cited
Batuwita S., de Silva M. and Edrisinghe U., 2013. A review of the danionine genera
Rasboroides and Horadandia (Pisces: Cyprinidae), with description of a new
species from Sri Lanka Ichthyol. Exploe. Freshwaters, Vol. 24, No. 2, 121-140
pp.
Chamikara S., 2012. පාට වර්ණ කොහි ගියාද හල්මල් දන්ඩියනේ? - සදාහරිත වෙළුම 1 කලාපය 9
Deraniyagala, P.E.P.1958. Three new cyprinoids, a new catfish
and variation among some cyprinoids and an anabantoid of Ceylon. Spolia
Zeylanica, 28: 129-138.
Meinken, H. 1957. Rabora nigromarginata n. spec. die
"Rote rasbora". Die Aquarien - und Terrarien Zeitschrift, 10: 65-68.
Pethiyagoda, R. 1991. Freshwater fishes of Sri Lanka.
Wildlife Heritage Trust, Colombo, xiii+362pp.